A team led by Oxford biologist Matilda Brindle has been honored with an Ig Nobel Prize. They earned the biomechanics award for establishing a formal definition of kissing, describing it as friendly, mouth-to-mouth contact between members of the same species that involves lip movement but excludes the exchange of food. This classification applies to various animals, including primates, polar bears, birds, and ants. According to their research, the behavior likely emerged in the ancestors of large apes between 16.9 and 21.5 million years ago, and it is highly probable that Neanderthals also engaged in kissing.

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